I have the following setup in many of my repositories:
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doc/README is a document with a vim:ft=rst modeline. As expected, this is rendered by GitHub as reStructuredText.
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Top-level README is a symlink to doc/README. This used to rendered by GitHub as reStructuredText until recently. But now it is shown as plain text. :-(
Examples:
I guess it might be an unintended consequence of fixing #1133?